Bosnian Croat war crimes suspect Miroslav Bralo has pleaded not guilty to 21 counts of war crimes and crimes against humanity during the 1993 Muslim-Croat fighting in Bosnia-Herzegovina.

Mr. Bralo appeared before United Nations war crimes tribunal in The Hague to answer such charges as murder, rape, torture, and illegal detention of civilians.

The indictment says Mr. Bralo committed the crimes as a member of a Bosnian Croat special unit called "The Jokers" that operated against Muslim forcers in central Bosnia.

He voluntarily surrendered to authorities last month.

Some information for this story provided by Reuters and AFP.