Croatian General Ante Gotovina pleaded not guilty Tuesday to four new charges of war crimes and crimes against humanity linked to the murders of Croatian Serbs in 1995.

Prosecutors at the Yugoslav war crimes tribunal in the Hague have charged Gotovina and two other Croatian officers, Ivan Cermak and Mladen Markac, with the crimes.

Gotovina was arrested a year ago on Spain's Canary Islands and remains in custody. He had pleaded not guilty to earlier charges.

Cermak and Markac have been released to await their trial. They appeared at Tuesday's hearing via a video link, and also pleaded not guilty to new charges.

The Hague war crimes tribunal indicted General Gotovina in 2001 for his role in the deaths of civilians during a 1995 offensive through a Serb-held area of the country.

Some information for this report was provided by AP and Reuters.