Russian bombers have test-fired missiles in the Bay of Biscay, between France and Spain, as part of air and naval exercises in the area.

The missiles were fired by two long-range Russian bombers Tuesday.

A Russian air force spokesman, Colonel Alexander Drobyshevsky, told the state news agency Itar-Tass that the flights are in strict accordance with international rules for using air space over neutral waters. 

Russia resumed long-range bomber flights in the North Atlantic last September.

The former Soviet Union regularly conducted such flights, but the practice ended when the Soviet Union collapsed.

Some information for this report was provided by AFP, AP and Reuters.